I'll preface this with "it happened in the 1980s" when I worked for Isuzu Trucks.
We had to do some fuel economy testing of a 4 tonne 4x4 truck that we hoped to sell to a rural fire brigades. Part of the testing included running the vehicle in 4x4 on dirt tracks and through a quarry for about half a shift, the other part of the testing was on a banked "speed loop" with the hubs in the free position and the locked position whilst in 4x2. The vehicle was fitted with fuel consumption gauges with a cross check by dipping/temperature check the tank before and after the tests and allowing the fuel to cool to the before test temp.
There was a marked difference between locked and free hubs - the figures escape me. When we won the contract I remember developing the driver training package which included having the hubs in free before driving on roads with a good surface - dirt or bitumen. The fuel consumption figures were significant enough that if they had to drive 100kms each way to a fire and then drive for six hours in rough terrain, they were pushing the limits of the vehicles safe fuel range.
I think there was also some engineering issues about the transfer case being "driven" by the front axle and not the engine. Probably no interaxle diff back in those days.
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